Leanchoilia illecebrosa

迷人林乔利虫

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分类学

门: 节肢动物门

纲: 大附肢纲

目: 林乔利虫目

科: 林乔利虫科

属:林乔利虫属

种: 迷人林乔利虫

描述: 

       迷人林乔利虫的身体分为头甲、躯干和尾节三部分。标本从20mm到46mm不等。头甲的长度约占身体总长(自头甲最前端至尾节末端,不包括大附肢和尾节的刚毛)的五分之一,尾节约为身体总长的六分之一。头甲之下有一对柄状眼和四对附肢,其中最前面的一对为大附肢,其余三对为双肢型附肢,与躯干部分的其他双肢型附肢形态类似。头甲最前部轻微向腹部弯曲,中后部至末端相对较直,最后稍微向上突起。头甲前边缘具有一个尖的喙状突起,头甲两侧的腹边缘较窄,头甲腹侧具有深色边缘线。具有一对复眼,其几乎全部覆盖在头甲之下,复眼与头甲之间的区域为眼睛的柄状构造。大附肢茎部由两个肢节组成,长度相对较大,能将远端的肢节向前或向下伸出一定的距离。远端有三个肢节,其中第一和第二远端肢节具有长的内突,内突末端具有长的鞭节;第三远端肢节与第一、第二远端肢节的内突齐长,末端具有一个爪刺以及一个鞭节。躯干具有11个体节,每个体节具有一对双肢型附肢。11个体节(背板)的长度差异不明显,尤其是第11个体节和前面的几个体节长度几乎相当,最宽的体节为第二、第三体节。背甲的侧边缘保存有锯齿状构造(头甲边缘没有)。躯干的附肢分为内肢和外肢,二者共同连接在基节上,基节从后侧看呈三角形,其内侧具有四个强壮的刺。外肢有两节,末端一节成椭圆形,具有长而密集的刚毛,刚毛成针状。内肢具有九个肢节,末端具有爪刺。尾节分为前端的似体节构造和后端的匕首状构造。匕首状构造的前端的基部呈柱状,两侧具有两条呈八字形的斜向的脊;后端呈倒三角形,比前端的基部短,两侧具有小而短的刚毛。

Taxonomy

Phylum: Arthropoda

Class: Megacheira

Order:Leanchoiliida

Family: Leanchoiliidae

Genus:Leanchoilia

Specie: Leanchoilia illecebrosa

Description:

Leanchoilia illecebrosa has a strongly convex dorsal exoskeleton consisting of a simple shield with a rostrum,11 trunk tergites and a pointed tailpiece with a marginal fringe of spines. The average size ranges from 20 to 46mm in length.  The shield covers the head,which consists of an anterior portion carrying the eyes and four limb-bearing segments. The eyes are located ventrally in the anterior region of the cephalon.They are distinctly reniform, or form part of a reniform structure. The first paired appendage in the cephalon is a prominent ʻgreat appendageʼ. The cephalon also carries three pairs of biramous appendages resembling those of the trunk.The ʻgreat appendageʼ consists of 6 articles. The two proximal articles do not show any outgrowths and are therefore regarded as forming a peduncle. The comparatively long third distal article has a hook-like extension medio-distally. The fourth distal article extends into another very long distal filament. The three long filaments were somewhat curved, and each of them tapers slowly in width toward the tip, indicating that they were flexible,but annuli are not discernible. In lateral aspect, the antero-lateral margins of the shield run ventrally in an almost straight horizontal line and then sharply flex dorsally straight up to meet medio-distally. In profile, the dorsal midline inclines gently to run almost straight back in its last quarter. In all, the lateral margins extend only slightly beyond the level of the venter. The anterior tip of the cephalic shield is drawn out into a short and straight rostrum. All the 11 trunk tergites of L. illecebrosa are almost of the same length, and the widest part is at the third trunk segment. The limbs posterior to the ʻgreat appendageʼ consist of a proximal portion, the basipod, on which an endopod with nine podomeres inserts distally, and an exopod with two portions inserts on its tilting outer margin. The limbs insert on the body proper through an ample membrane. The tailpiece consists of a cylindrical part anteriorly and a dagger-shaped flap posteriorly. Each side of the tail flap has a row of postero-laterally pointed needle-like setae.

年龄和产地

时代:寒武系第二统第三阶,始莱德利基虫-武定虫化石层

主要产地: 中国云南澄江生物群(筇竹寺组玉案山段)

Age & Localities

Age:Cambrian Series 2, Stage 3, Eoredlichia-Wutingaspis Zone

Principal localities: Chengjiang biota(Yu’anshan Member of the Chiungchussu Formation),eastern Yunnan Province, China

Reference:

Yu Liu, Xian-Guang Hou & Jan Bergström,2007.Chengjiang arthropod Leanchoilia illecebrosa (Hou,1987) reconsidered.GFF,Volume 129,Issue 3.

Yu Liu,2014.A 520 million-year-old chelicerate larva.Nature Communications volume 5, Article number: 4440.

Joachim T Haug, Derek EG Briggs and Carolin Haug,2012.Morphology and function in the Cambrian Burgess Shale megacheiran arthropod Leanchoilia superlata and the application of a descriptive matrix.BMC Evolutionary Biology volume 12, Article number: 162.

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